The DENNISON DNA Project: Patrilineage 2

DENNISON DNA Patrilineage 2

The three American DENNISONs appear to have emanated from Pittsylvania County, Virginia, in the late 1700s. Even though Pittsylvania lies just to the SE of the Blue Ridge mountains, just outside the Valley of Virginia where the Scotch-Irish DENNISONs of Patrilineage 1 settled, the yDNA results shows that these two groups of Virginia DENNISONs are completely unrelated.

It is highly unlikely that the Patrlineage 2 DENNISONs are Scotch-Irish, for a number of reasons, beginning with the fact that most British DENNISONs are English, not Scottish, thus the vast majority of American DENNISONs have English, not Scotch-Irish roots.

Second, Pittsylvania County was overwhelmingly settled, not by the Scotch-Irish whose initial settlements were beyond the mountains, but by tidewater Virginians slowly trickling west. This was true of Pittsylvania's parent county, Halifax, to the east, and of its child county, Henry, to the west. And the same east to west pattern applied to the next northern tier of counties, from Lunenburg, to Bedford, to Franklin County (where I happen to live). Furthermore, these early settlers of Virginia’s piedmont, and “south side”, continued west mostly in the wake of the Scotch-Irish, progressing to Tennessee and Kentucky a decade or two later.

Third, the early settlers of these cismontane counties, including these DENNISONs, do not, by and large, bear Scotch-Irish surnames, nor are their given names confined to the restrictive Scotch-Irish set, and they don’t appear to observe the customary Scottish and Scotch-Irish onomastic (child-naming) pattern either.

Positive yDNA Evidence of an English Origin

Finally, we have a first piece of DNA evidence that England is the original home of this early Virginia family of DENNISONs, in the form of a new member of Patrilineage 2, Lee Daniels—a Londoner whose ancestors have always been English back at least to Thomas Daniel of Buckinghamshire, born say 1762. This evidence is provisional, though, for two reasons. First, and most obvious, is the different surname, and second is the fact that there is a genetic distance of 6 between Lee and the DENNISON-surnamed members of this patrilineage.

As for the surname, experienced FTDNA project administrators are finding ample confirmation in their data, of the well-known phenomenon of the MPE (Misattributed Paternity Event), where a son ends up with a different surname from his biological father. The rate of MPEs per generation in historical western societies is variously estimated at between 1-7%, depending on the place and period, and since the probabilities are cumulative, at a rate of 3%, the chances of an MPE over, say, 8 generations, would be about 20%. Not all MPEs result in a changed surname, but if we guess that this is true of half of them, we might expect to find one divergent surname for every 10-20 in a yDNA surname project, and that is about what project administrators are finding in their data.

Moreover, it’s probable that many patrilineage members with divergent surnames are being overlooked—stranded in the projects whose surname they match—first, and understandably, because we all have a considerable investment in the assumption that the surname we bear represents the ancestral “tribe” we belong to, and second, thanks to an unfortunate set of genetic distance guidelines published by FTDNA (the premier yDNA testing company), which have misled many, if not most, project administrators.

According to these guidelines, two living men tested on FTDNA’s 37-marker panel whose results come in at a genetic distance of 6 or more, are “unrelated”, although for a GD of 6, it is allowed that a person might be considered a member of the patrilineage if a “tweener” can be found. What is meant by this, is that if an additional testee can be found who is GD 5 or less with both the accepted members and with the person in question, that the latter may thereby be linked with the group.

But these guidelines fail to take into account either the fact that two people who match only loosely also have a common surname (in most cases), and furthermore, that in many cases there is strong genealogical evidence of a relationship. And when these factors are taken into account, examples of people who belong in patrilineages even though they have at best a GD 8-10 relationshp with the others, are not hard to find. The fact is, there are many cases where as many as 5 mutations have occurred in lines of descent since the founding of a surname line, which may go back as many as 800-900 years, and if there happen to be two such lines of descent within the same patrilineage, each with 5 mutations different from the other, then even a GD of 10 is possible, although to be sure one would certainly want to find a “tweener” for such a tenuous relationship.

In the instant case, the GD is only 6—well within the bounds of probability, requiring only that there have been two lines of descent since the founder of DENNISON Patrilineage 2, each with 3 different mutations. Furthermore, we have, as further confirmation, a 67-marker comparison between Lee (D-11) and Dale Dennison (D-10), with just one additional marker divergence. Although in the TMRCA chart below, I have calculated that there is a 50% probability that these two have a common ancestor within the last 510 years, the FTDNA Tip calculator lowers this to 460 years, even after factoring in that we know there was no common ancestor since 1750. And there is also the consideration that Lee Daniels is completely unrelated to any of the 54 other members of the DANIEL(S) surname project who have been tested on 37 markers or more and who share the usual British haplogroup R1b.

Although a tweener would be desirable in this case, especially since the surname is different, in my judgement the odds greatly favor the proposition that Lee Daniels belongs to the same patrilineage as the existing DENNISON members. As I say, there are any number of such instances to be found in the 5000+ FTDNA surname projects, but I have yet to hear of an authenticated case of a false positive, where a person with a different surname and a reasonably close GD can be shown by historical research not to belong to a patrilineage—although I am sure that such cases exist. Let us by all means keep our minds open to both alternatives here, as we wait for a suitable tweener to emerge to confirm the relationship beyond any reasonable doubt.

Finally, if, as I think, Lee Daniels belongs with this group, we also have to recognize the possibility that the original surname for this group was DANIEL(S), not DENNISON. All we really have here is evidence of a probable close relationship between Lee and a single rather closely related line of DENNISONs. We don’t as yet have any history on either ancestry before the middle of the 18th century that would give us some basis for choosing one surname as the original one over the other. However, the fact that Lee matches to none of the 54 other members of the DANIEL(S) project does suggest that the original surname was DENNISON. If a match should turn up between Lee and a newly tested DANIELS, it may or may not tip the balance toward DANIEL(S) as the original surname, or for the inclusion of Lee in the DENNISON group; for example, the new DANIELS might prove to be of the same descent as Lee—from Thomas Daniel of Buckinghamshire.

DENNISON Patrilineage 2 Descendancies

The following ancestral DENNISON descendancies have been contributed by researchers of Patrilineage 2. Each descendancy begins with the earliest known male ancestor of a particular sub-lineage and continues down to the tested male descendant. Since the DNA Surname Project is focused on tested or testable male DENNISON lines, these descendancy trees have been pruned not only of daughters, but also of male lines that have “daughtered out”. However, complete reconstructed families of the first generation or two will be included because of their broad-based genealogical interest.

The information provided for each male DENNISON should be sufficient, in most cases, to uniquely identify him in the USCensus and other readily available sources. These data comprise (insofar as is known): date and place of birth, date and place of death, and the name(s) of his marriage partner(s). Indefinite dates are always qualified as “abt” or “say”. Dates given as “abt” imply supporting evidence that merely fails of complete accuracy, while “say” dates are guesstimates based on typical patterns of the time, place, and social group.

The yDNA-tested male descendants are flagged below with their Project#s, e.g.  D-08).

invisible writing

1-William Dennison of Washington County, Virginia, born abt 1783

(source: Julie Dennison)

1--William Dennison (abt 1783 PittsylvaniaCoVA - aft 12Oct1860)  m. Jane Barnes

|--2-Trammel Dennison (abt 1807 VA - 1891 GraysonCoKY)  m. Frances Carter

|   |--3-Henry Clay Dennison (1832 GraysonCoKY - 1890 GraysonCoKY)

|   |   ---m. Elizabeth A. Weedman

|   |   |--4-Charles William Dennison (1857 KY - 1939 SangamonCoIL)  m2. Sarah T. Kiper

|   |   |   |--5-Clarence E. Dennison (1895 GraysonCoKY - 1965 McLeanCoIL)

|   |   |   |   ---m. Elizabeth Cotton

|   |   |   |   |--6-William Lee Dennison (1944 McLeanCoIL - 2001 IL)

|   |   |   |   |   |--7-son of William  *** D-08 ***

invisible writing

1-Joel Dennison of Virginia, born abt 1808

(source: Lois Franceschi)

1--Joel Dennison (abt 1808 VA -)  m. Anna Jackson

|--2-James W. Dennison (abt 1839 KY -)

|--2-John Burch Dennison (1840 KY - 1917 ReynoldsCoMO)

|--2-Daniel Albert Dennison (1844 KY - 1912)

|--2-Jeremiah M. Dennison (1849 MO - 1920 IronCoMO)

|--2-Benjamin Franklin Dennison (1852 ReynoldsCoMO - 1934 ReynoldsCoMO)

|   ---m. Rebecca Jane Stevens

|   |--3-Elmer Joel Dennison (abt 1905 MO -)  m. Virgie Marie Henson

|   |   |--4-James Dale Dennison  m. Irma Sue Rood  *** D-10 ***

invisible writing

1-Thomas Daniel of Buckinghamshire, England, born say 1762

(source: Lee Daniels)

1--Thomas Daniel (say 1762 Bucks ENG - 1828 Bucks ENG)  m. Judith Stears

|--2-Joseph Daniels (1805 Bucks ENG -)  m. Sarah Tims

|   |--3-William Daniels (1830 Bucks ENG -)  m. Hannah Braggins

|   |   |--4-Andrew William Daniels (1865 Bucks ENG -)  m. Lavinia Jones

|   |   |   |--5-Ernest John Daniels (1897 London ENG - 1951)  m. Dorothy Caroline Watts

|   |   |   |   |--6-John Frederick Daniels (1943 London ENG -)  m. Joan Ann Winslade

|   |   |   |   |   |--7-Lee John Daniels (1966 London ENG -) *** D-11 ***

|--2-Edward Daniels (1807 -)

|--2-George Daniels (1808 - 1887)

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Navigating from here

The menu buttons at top right take you to other pages on this site, while the nav panel above targets other points on this page, or brings up other resources (papers I’ve written, and the like). If you find yourself lost, the browser BACK button will take you back to where you were (some people also have a convenient BACK button on their mouse, right under their thumb). Or hitting the HOME key of your keyboard will take you back to the top of this page where you are now.

Some Key Terms: haplotype, haplogroup, patrilineage, RPH.

DENNISON Patrilineage 2 Researchers

Active researchers of this DENNISON patrilineage are shown below. Those with highlighted names may be e-mailed by clicking on their names, and their posted descendancies may be viewed by clicking on their highlighted Patrilineage Project#s. Satellite members of the project are listed immediately after the principal researcher for the sublineage they are interested in, and their names are preceded by a dash. Where the person tested is not also the principal researcher, the former’s name appears under the latter’s, in parentheses.

Proj# Researcher
(Test Subject)
Test
Panel
FTDNA
Kit# 
YSEARCH
ID
D-08 Julie Dennison F37/A43 168547
D-09 Bob Dennison F37 96122 2NA8M
D-10 Lois Franceschi
(Dale Dennison)
F67 151490
D-11 Lee Daniels F67 13863 8JB52

The Deep Lineage of Haplogroup R-L21+

A man’s yDNA may be classified according to a deeper, broader, ancestral schema by testing ySNPs instead of ySTR markers. These ySNPs are unique point mutations to the yChromosome that occur so much less frequently than ySTR mutations that they are of no use in sorting people into patrilineages, but by the same token they are ideal for sorting patrilineages into the various haplogroups and subclades of the broad human population. And by correlating the haplogroup subclades of both modern descendants and of their ancient ancestors (by testing their remains) with geographic population flows, and with archaeological evidence signifying cultural groupings, the whole broad history of homo sapiens is gradually being reconstructed.

DENNISON Patrilineage 2 falls into haplogroup R1b1b2, and with the addition of Lee Daniels to the project, who has recently done deep clade testing, he and all other members of the group can now be more specifically classified as subclade R1b1b2a1b5—or R-L21+ in the new nomenclature that identifies subclades by their most recently differentiated SNP mutation. Each new mutation represents a further branching of the human tree, defining a new subclade. The “+” means that additional SNPs have been identified since Lee tested negative on M222, namely L193, L226, M37, and P66, and it is possible (in this case likely) that further SNP testing would show that DENNISON Patrilineage 2 would follow one of the branching sublineages of L21.

As it is, the L21+ subclade of R1b is very close to the Western Atlantic Modal Haplotype (WAMH) that is found all along the European seaboard from Portugal to Scandinavia, and R1b is by far the most common subclade in the British Isles. The L21 SNP mutation is thought to have originated in southern or Alpine Germany about 4000 years ago, and spread north and west from there into the European continent, and thence to Britain, and especially Ireland and Scotland, where, like the Celtic culture, it predominates. However, it's a mistake to identify L21, per se, with the Celtic, or Hallstatt culture that it long predates. Here is a map showing the spreading out and differentiation of R1b over the last 20,000 years. In a new book (published May2011), “The Scots: A Genetic Journey”, by Alistair Moffat and Jim Wilson, the authors have identified the SNP S145, which has always been found so far associated with L21, as a marker for the ancient Pictish culture of Scotland, but as the R1b distribution map indicates, L21 is widely distributed throughout Britain, and takes it’s origins many thousands of years ago on the NW European continent.

At the rate progress is being made in the reconstruction of human history through DNA analysis, before too many more years have passed, I expect that some of these subclades will be brought down into genealogical time, and may even become a shortcut means of identifying patrilineages. In the meantime, you can read more about haplogroups and their distribution across the continents at this site, and there are already efforts to further break down the broad L-21* group by means of haplotype analysis on this DNA Forum thread (you need to register with the Forum, though, to access it).


DENNISON Patrilineage 2 yDNA Haplotypes Compared

These charts provide some idea of the closeness of relationship between each pair of test subjects of this patrilineage. The cell at the intersection of each column/row pair shows either the GD (Genetic Distance) between the pair (basically, the number of mutations), or the estimated TMRCA—the Time in years back to the Most Recent Common Ancestor of the pair (not the MRCA of the whole patrilineage). For an extended discussion of the application of these concepts, click here.

While the GD is exact, there is no obvious way to tell how the mutations divide between the two subjects, because the haplotype of their common ancestor, from whom they have mutated, is unknown. However, I have developed a procedure using the GD chart for inferring the haplotype of the common ancestor, which I call the Root Prototype Haplotype (RPH), and I use this as a basis for marking mutations in the yDNA Haplotypes chart, following. The FTDNA commentary says that relationships are only possible where the GD between two subjects is less than 6, but this is a mistake. A GD of 6 from the common ancestor would push that ancestor back before the origin of most surnames, but because GD is the sum of the mutations down each of two lines of descent from the founder, there might be no more than 3 mutations to each haplotype which is well within the relationship guidelines. In fact, a GD of up to 10 between two project haplotypes is conceivable, as long as they bear a common surname.

TMRCA provides at best a very loose estimate of the time back to the common ancestor of two patrilineal descendants, so I have restricted the following set of estimates for paired members of this project just to the members who have tested out to 67 markers. The time may be measured in generations or years, but I find the year estimate more useful. I use 34 for the number of years per generation, based on a number of published studies, as well as on an informal one of my own, and I've found that the best way to project back to an earlier ancestor is to substract the TMRCA estimate from 1950 (a notional birth year for the a typical contemporary testee) to obtain an estimate for the birth date of the MRCA of each pair of subjects.

In the TMRCA estimates below there is an equal probability that the MRCAncestor of each pair of subjects was born earlier or later than the projected date; indeed, he could quite easily have been born 100 years or more earlier. Besides the inherent inaccuracy of TMRCA estimates (given the limitations of our present scientific knowledge of the mutation process), the values in this TMRCA chart need to be further adjusted to account for the fact that it is known that none of the current members are related to each other within the last 4 generations. This pushes the TMRCA estimates back another 50-75 years. The FTDNA Tip calculator (available from each project member’s personal FTDNA page) is able to factor in such knowledge, but Tip is currently limited to making comparisons between just two individuals at a time. The following chart is meant to provide a rough overview of the closeness of the genetic relationships between members within a time framework. There is much else which can be said about TMRCA, and in fact I have had my say here.

37-Marker Haplotype Comparison Matrices (low numbers are closest)

Genetic Distances, DENNISON Patrilineage 2, from 37-marker yDNA Comparisons TMRCAs, DENNISON Patrilineage 2 Patrilineage, from 37-marker yDNA Comparisons

The number in each cell is the number of divergent mutations
between each pair of haplotypes.

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What FTDNA has to say about Genetic Distance for 37-marker comparisons

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67-Marker Haplotype Comparison Matrices (low numbers are closest)

Genetic Distances, DENNISON Patrilineage 2 Patrilineage, from 67-marker yDNA Comparisons TMRCAs, DENNISON Patrilineage 2 Patrilineage, from 67-marker yDNA Comparisons

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The number in each cell is the number of divergent mutations
between each pair of haplotypes.

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What FTDNA has to say about Genetic Distance for 67-marker comparisons


DENNISON Patrilineage 2 yDNA Haplotypes

The chart below shows the haplotypes for each tested project member of this patrilineage. I've decapitated most of the marker names (truncating “DYS393” to just “393”) to improve readability. The colored markers mutate slower or faster than the norm. Thus, [DYS]439 is fast, [DYS]458 is faster, and CDYa&b are blazing, while [DYS]393 is slow. Contrary to what one might think, though, it makes very little difference to the TMRCA calculation whether the markers that mutate are slow or fast. One expects most of the mutations to occur amidst the fast markers, and if slow markers mutate instead that actually increases the TMRCA a bit.

Test Subject Information FTDNA 37-Marker Panel FTDNA Markers 38-67 Additional Markers

Proj
#
 Principal
 Researcher
Earliest Known
DENNISON Ancestor

Name, BirthYear,
BirthPlace
3
9
3
3
9
0
1
9
/
3
9
4
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
I
3
9
2
3
8
9
I
I
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
4
6
0
Y
G
A
T
A
H
4
Y
C
A
I
I
a
Y
C
A
I
I
b
4
5
6
6
0
7
5
7
6
5
7
0
C
D
Y
a
C
D
Y
b
4
4
2
4
3
8
5
3
1
5
7
8
3
9
5
S
1
a
3
9
5
S
1
b
5
9
0
5
3
7
6
4
1
4
7
2
4
0
6
S
1
5
1
1
4
2
5
4
1
3
a
4
1
3
b
5
5
7
5
9
4
4
3
6
4
9
0
5
3
4
4
5
0
4
4
4
4
8
1
5
2
0
4
4
6
6
1
7
5
6
8
4
8
7
5
7
2
6
4
0
4
9
2
5
6
5
4
4
1
4
4
5
4
5
2
4
6
1
4
6
2
4
6
3
6
3
5
G
A
A
T
1
B
0
7
Y
G
A
T
A
A
1
0
D-08  Julie Dennison WilliamH-b.c1783 VA 13 23 14 10 11 14 12 12 12 13 12 29 17 9 10 11 11 24 15 19 33 15 15 17 17 10 11 19 23 17 15 19 18 37 39 12 12 12 14 141231121124915
D-09  Bob Dennison Stephen-b.c1789 VA 13 24 14 10 11 14 12 12 12 13 12 29 17 9 10 11 11 24 15 19 33 15 15 17 17 10 11 19 23 17 15 19 18 37 38 12 12
D-10  Lois Franceschi Joel-b.c1808 VA 13 24 14 10 11 14 12 12 12 13 12 29 17 9 10 11 11 24 15 19 32 15 15 17 17 10 11 19 23 17 15 19 18 37 38 12 12 11 9 15 16 8 11 10 8 10 10 12 23 23 16 10 12 12 15 8 12 21 21 14 12 11 13 11 11 12 12
D-11  Lee Daniels ThomasDaniels-b.c1762 13 24 14 10 11 14 12 12 12 13 13 29 17 9 10 11 11 24 15 19 30 14 15 15 17 10 12 19 23 17 15 20 18 37 38 12 12 11 9 15 16 8 11 10 8 10 10 12 23 23 16 10 12 12 15 8 12 22 21 14 12 11 13 11 11 12 12

You may click on highlighted Project#s (like D-08) to see the posted pedigree for a particular test subject. Click on highlighted Researcher names, like Alan Denison to go to the project directory that shows the full names of the members, and provides clickable e-mail links for the names highlighted.

The test subject whose “Earliest Known Ancestor” is colored red is the one whose haplotype differs the least from all the others and is therefore designated the Root Prototype Haplotype (RPH)—the haplotype that is likely to be the closest to that of the Most Recent Common Ancestor (MRCA) of the group. Marker values that deviate from those of the RPH are deemed to be mutations, and are highlighted in lime green—or tomato, for multistep mutations—two or more separate mutations to the same marker.

Where multicopy markers DYS464 and YCA (each taken as a whole) diverge in value from those of the RPH, the whole adjacent set of values will be colored yellow green, and will be counted as a single mutation. In the same way, reclOH mutations, which may affect several blocks of separated markers, will be colored orange and treated all as a single mutation for purposes of calculating Genetic Distance. There is a small probability in both cases that more than one mutation to the set has occurred, and where this is strongly enough suspected, part of the block(s) may be colored tomato. like other multistep mutations.

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